If held by a Heracross, the Heracronite allows it to Mega Evolve into Mega Heracross if its Trainer has a Key Stone. A Trainer can only Mega Evolve one Pokémon per battle (or only have one Mega-Evolved Pokémon at any given momentZA).
As a Mega Stone, it is unaffected by item-negating effects (such as Embargo and Klutz). Item-manipulating effects (such as Trick and Fling) cannot give or remove a Heracronite from Heracross.
Outside of battle
If held by a Heracross, the Heracronite allows it to Mega Evolve into Mega Heracross when it performs a Spectacular Talent during a Pokémon Contest Spectacular if its Trainer has a Key Stone. Heracross will remain a Mega Heracross until the Talent Round ends.
Adds one more Mega Heracross to the left of the match. Actually converts the tiles directly to the left of all other Mega Heracross into a Mega Heracross. This effect can replace any disruption in the tile above and counts as erasing the content of the tile for damage, but does not affect black-cloud disruptions. Does nothing if the left tile is empty.